Human Inclusive Fitness will measure the success of the Colorful Prince's mission.
Unlike environments and social programs that can be assessed with post-occupancy or Person–Environment Fit evaluations––the degree to which an individual or group’s biological or psychological needs, values, goals, abilities, job, or role match the constructed landscape––the metric for assessing the success of niche construction and gene-culture coevolution habitat is the number of offspring that an individual rears, and supports regardless of whether a blood relationship exist or not.
Niche constructions are safe spaces where caregivers with or without blood relation to children embrace the task of childrearing, sharing the parenting role, and offering multiple sources of attachment––mirroring the skinship structural reciprocity model.
From a gene-evolution perspective, success is measured as personal fitness, the number of children one can produce. Human inclusive fitness, however, implies unconditional emotional attachment, an intrinsic need to nurture, and social cooperation, all of which are altruistic values.
With inclusive fitness exercised in this manner, children receive enhanced security and an improved capacity for viewing the world from multiple perspectives. As a component of nurturing, they gain a variety of approaches to imitate that consistently carry a bias favoring altruism, the concern for the well-being and happiness of others above oneself.
In addition, the social and physical structure of the niche also conveys a context bias, suggesting that cooperative altruistic behavior is normal, and the implied content bias recommends that it be adopted.